From Lylat Wiki, your source on Star Fox information. By Fans, for Fans.
Zako is a shark-like alien and a bounty hunter hired to work for the Anglar Empire. He appears in Star Fox Command.
Zako appears to be under the command of Octoman, as he was ordered to kill Slippy and Amanda. Both Zako and Octoman appear as enemies in the planet Aquas, where Zako pilots the Devil Shark. In some of the other paths, he guards a base on the last mission in the acidic Venom sea, but ends up being killed.
Zako's name could be a pun on the Japanese word "zako", meaning "smallfry".
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